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We have seen that the neutral forms of radiation, photons and neutrons, are very penetrating and can pass through layers of material without interacting In these cases, we need to distinguish between the amount of radiation exposure and the amount of energy absorbed by the material Photons such as X-rays, bremsstrahlung, and g rays play an important role in nuclear medicine, but they are not strongly absorbed by tissue The exposure to these photons is not equal to the dose In the case of highly ionizing radiation such as charged particles, the exposure will correspond to the absorbed energy except for very thin materials that allow the particle to escape In order to characterize radiation effects, we need to know the amount of energy absorbed by the material, which is called the absorbed dose The unit of radiation exposure is the roentgen (R).

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It is a historical unit of the exposure and characterizes the radiation incident on an absorbing material without regard to the character of the absorber The unit was formalized in 1928 as The amount of radiation which produces one electrostatic unit of ions (esu), either positive or negative, per cubic centimeter of air at standard temperature and pressure Translated in modern units: 1 roentgen 2:58 10 4 coulomb=kg-air 0:3336 nC=cm3 at STP This value corresponds to an absorbed energy of approximately 88 mJ/kg using the effective ionization energy of 34 eV per ion pair in air The roentgen is most often used to describe the intensity of a photon source such as a medical X-ray machine or other irradiator The exposure should be measured at some distance from the source so that the radiation eld is uniform compared to the dimensions of the detector.

The detector is usually an ion chamber lled with dry air that is sensitive to pico-coulombs of charge As studies of the effects of exposure to all types of radiation went on, it became clear that these effects were correlated with the amount of absorbed energy, which is generally less than the exposure In 1962 the rad was formally de ned as a special unit of energy called the radiation absorbed dose with a value of 100 ergs per gram of absorbing material The rad is a convenient physical standard that correlates well.

The major problem in measuring reactions consists in selecting small absorption bands of molecular groups which undergo reactions from the large background absorption of water and of the entire protein. This difficulty is met by obtaining difference spectra between spectra of the protein in its ground state and in an activated state. Such measurements require highly sensitive instrumentation. FTIR has two advantages over conventional dispersive IR spectroscopy, i.e., the multiplex and the Jaquinot advantage. This makes it possible to reliably detect such small absorption changes with a time resolution of a few nanoseconds. Alternatively, if it is sufficient to measure only absorption changes within a bandwith of a few wavenumbers, photolysis setups with dispersive elements providing a time resolution down to picoseconds can be used. Instead of presenting a comprehensive literature review, this section develops the basic ideas of FTIR reaction mechanism studies. Examples include the light-induced proton-pump mechanism of the membrane protein bacteriorhodopsin and the light-induced electron transfer reactions of bacterial photosynthetic reaction centers. For detailed reviews, see (Braiman and Rothschild, 1988; Kitagawa and Maeda, 1989; Gerwert, 1993).

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